Skin Cutaneous Melanoma_SKCM_TCGA-FS-A1ZM-06A-12R-A18S-07 Gene Set

Dataset TCGA Signatures of Differentially Expressed Genes for Tumors
Category transcriptomics
Type tissue sample
Description tissue sample derived from Skin Cutaneous Melanoma_SKCM (The Cancer Genome Atlas)
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Genes

437 genes with high or low expression in Skin Cutaneous Melanoma_SKCM_TCGA-FS-A1ZM-06A-12R-A18S-07 relative to other tissue samples from the TCGA Signatures of Differentially Expressed Genes for Tumors dataset.

high expression

Symbol Name
A2M alpha-2-macroglobulin
ABCB4 ATP-binding cassette, sub-family B (MDR/TAP), member 4
ACSM3 acyl-CoA synthetase medium-chain family member 3
ADAMTS3 ADAM metallopeptidase with thrombospondin type 1 motif, 3
ADCY1 adenylate cyclase 1 (brain)
ADCY3 adenylate cyclase 3
AEBP2 AE binding protein 2
AFF1 AF4/FMR2 family, member 1
AIM2 absent in melanoma 2
ALG10 ALG10, alpha-1,2-glucosyltransferase
ALG10B ALG10B, alpha-1,2-glucosyltransferase
ANK2 ankyrin 2, neuronal
ANKRD17 ankyrin repeat domain 17
APLF aprataxin and PNKP like factor
ARHGEF28 Rho guanine nucleotide exchange factor (GEF) 28
ARSB arylsulfatase B
ASAP2 ArfGAP with SH3 domain, ankyrin repeat and PH domain 2
ASXL2 additional sex combs like transcriptional regulator 2
ATN1 atrophin 1
ATP10D ATPase, class V, type 10D
ATP2B1 ATPase, Ca++ transporting, plasma membrane 1
ATP8B2 ATPase, aminophospholipid transporter, class I, type 8B, member 2
B3GNT2 UDP-GlcNAc:betaGal beta-1,3-N-acetylglucosaminyltransferase 2
BAALC brain and acute leukemia, cytoplasmic
BBS12 Bardet-Biedl syndrome 12
BICD1 bicaudal D homolog 1 (Drosophila)
BIRC6 baculoviral IAP repeat containing 6
BST1 bone marrow stromal cell antigen 1
BSX brain-specific homeobox
BTBD19 BTB (POZ) domain containing 19
C12ORF4 chromosome 12 open reading frame 4
C12ORF66 chromosome 12 open reading frame 66
C12ORF71 chromosome 12 open reading frame 71
C12ORF73 chromosome 12 open reading frame 73
C22ORF31 chromosome 22 open reading frame 31
C2CD5 C2 calcium-dependent domain containing 5
C2ORF16 chromosome 2 open reading frame 16
C2ORF50 chromosome 2 open reading frame 50
C2ORF70 chromosome 2 open reading frame 70
C3ORF67 chromosome 3 open reading frame 67
C5ORF52 chromosome 5 open reading frame 52
C6ORF89 chromosome 6 open reading frame 89
C7ORF65 chromosome 7 open reading frame 65
CAND1 cullin-associated and neddylation-dissociated 1
CAPN14 calpain 14
CAPRIN2 caprin family member 2
CASD1 CAS1 domain containing 1
CASP5 caspase 5, apoptosis-related cysteine peptidase
CCDC121 coiled-coil domain containing 121
CCDC144A coiled-coil domain containing 144A
CCDC149 coiled-coil domain containing 149
CCDC167 coiled-coil domain containing 167
CCDC91 coiled-coil domain containing 91
CCKBR cholecystokinin B receptor
CCL3 chemokine (C-C motif) ligand 3
CCNG2 cyclin G2
CD9 CD9 molecule
CHRD chordin
CHRNA6 cholinergic receptor, nicotinic, alpha 6 (neuronal)
CLCC1 chloride channel CLIC-like 1
CLCN3 chloride channel, voltage-sensitive 3
CLIP4 CAP-GLY domain containing linker protein family, member 4
CMAS cytidine monophosphate N-acetylneuraminic acid synthetase
CMTR1 cap methyltransferase 1
COCH cochlin
COPS7A COP9 signalosome subunit 7A
COPZ2 coatomer protein complex, subunit zeta 2
COX18 COX18 cytochrome c oxidase assembly factor
CPEB2 cytoplasmic polyadenylation element binding protein 2
CPNE8 copine VIII
CREB5 cAMP responsive element binding protein 5
CREBL2 cAMP responsive element binding protein-like 2
CRISP3 cysteine-rich secretory protein 3
DDX11 DEAD/H (Asp-Glu-Ala-Asp/His) box helicase 11
DDX12P DEAD/H (Asp-Glu-Ala-Asp/His) box polypeptide 12, pseudogene
DDX47 DEAD (Asp-Glu-Ala-Asp) box polypeptide 47
DEFB109P1B defensin, beta 109, pseudogene 1B
DHX15 DEAH (Asp-Glu-Ala-His) box helicase 15
DHX29 DEAH (Asp-Glu-Ala-His) box polypeptide 29
DKFZP586I1420 uncharacterized protein DKFZp586I1420
DMRTA1 DMRT-like family A1
DNAJB14 DnaJ (Hsp40) homolog, subfamily B, member 14
DNAJC22 DnaJ (Hsp40) homolog, subfamily C, member 22
DNAJC27 DnaJ (Hsp40) homolog, subfamily C, member 27
DNM1L dynamin 1-like
DOCK4 dedicator of cytokinesis 4
DPY19L1 dpy-19-like 1 (C. elegans)
DSPP dentin sialophosphoprotein
DSTYK dual serine/threonine and tyrosine protein kinase
DUSP18 dual specificity phosphatase 18
DUSP19 dual specificity phosphatase 19
DYNAP dynactin associated protein
DYRK2 dual-specificity tyrosine-(Y)-phosphorylation regulated kinase 2
E2F6 E2F transcription factor 6
EFHC2 EF-hand domain (C-terminal) containing 2
EFR3B EFR3 homolog B (S. cerevisiae)
EIF4E eukaryotic translation initiation factor 4E
EMX2 empty spiracles homeobox 2
EMX2OS EMX2 opposite strand/antisense RNA
ENOPH1 enolase-phosphatase 1
EPB41L1 erythrocyte membrane protein band 4.1-like 1
EPDR1 ependymin related 1
EPT1 ethanolaminephosphotransferase 1
ERC1 ELKS/RAB6-interacting/CAST family member 1
ERGIC2 ERGIC and golgi 2
ETNK1 ethanolamine kinase 1
ETV6 ets variant 6
EVC2 Ellis van Creveld syndrome 2
EXOC1 exocyst complex component 1
FAM106CP family with sequence similarity 106, member C, pseudogene
FAM114A1 family with sequence similarity 114, member A1
FAM134B family with sequence similarity 134, member B
FAM184B family with sequence similarity 184, member B
FAM66C family with sequence similarity 66, member C
FBXL5 F-box and leucine-rich repeat protein 5
FBXO41 F-box protein 41
FBXO47 F-box protein 47
FGF2 fibroblast growth factor 2 (basic)
FGFR1OP2 FGFR1 oncogene partner 2
FHAD1 forkhead-associated (FHA) phosphopeptide binding domain 1
FKBP9 FK506 binding protein 9, 63 kDa
FKBP9P1 FK506 binding protein 9 pseudogene 1
FNDC4 fibronectin type III domain containing 4
FOXD4L5 forkhead box D4-like 5
FOXJ2 forkhead box J2
FOXL2NB FOXL2 neighbor
FRAS1 Fraser extracellular matrix complex subunit 1
FRYL FRY-like
GABARAPL1 GABA(A) receptor-associated protein like 1
GAL3ST4 galactose-3-O-sulfotransferase 4
GALNT7 polypeptide N-acetylgalactosaminyltransferase 7
GALNT8 polypeptide N-acetylgalactosaminyltransferase 8
GALR2 galanin receptor 2
GAS7 growth arrest-specific 7
GAS8-AS1 GAS8 antisense RNA 1
GDF11 growth differentiation factor 11
GFPT1 glutamine--fructose-6-phosphate transaminase 1
GLCCI1 glucocorticoid induced 1
GLI3 GLI family zinc finger 3
GLS2 glutaminase 2 (liver, mitochondrial)
GNAS GNAS complex locus
GNS glucosamine (N-acetyl)-6-sulfatase
GOLGA8CP golgin A8 family, member C, pseudogene
GOLT1B golgi transport 1B
GPBP1 GC-rich promoter binding protein 1
GPN1 GPN-loop GTPase 1
GPR162 G protein-coupled receptor 162
GPRASP1 G protein-coupled receptor associated sorting protein 1
GRK4 G protein-coupled receptor kinase 4
GSG1L GSG1-like
GTF3C2 general transcription factor IIIC, polypeptide 2, beta 110kDa
HAUS3 HAUS augmin-like complex, subunit 3
HCCAT5 hepatocellular carcinoma associated transcript 5 (non-protein coding)
HCG18 HLA complex group 18 (non-protein coding)
HCG9 HLA complex group 9 (non-protein coding)
HEATR5B HEAT repeat containing 5B
HECW1 HECT, C2 and WW domain containing E3 ubiquitin protein ligase 1
HEY1 hes-related family bHLH transcription factor with YRPW motif 1
HIST4H4 histone cluster 4, H4
HOXB13 homeobox B13
HOXC10 homeobox C10
HOXC11 homeobox C11
HOXC12 homeobox C12
HOXC9 homeobox C9
HSPA12A heat shock 70kDa protein 12A
ICA1 islet cell autoantigen 1, 69kDa
IFFO1 intermediate filament family orphan 1
IFNA5 interferon, alpha 5
IL27RA interleukin 27 receptor, alpha
IL6ST interleukin 6 signal transducer
INPP4A inositol polyphosphate-4-phosphatase, type I, 107kDa
IPO8 importin 8
ITGA2B integrin, alpha 2b (platelet glycoprotein IIb of IIb/IIIa complex, antigen CD41)
ITGB3 integrin, beta 3 (platelet glycoprotein IIIa, antigen CD61)
ITPR1 inositol 1,4,5-trisphosphate receptor, type 1
KCNH6 potassium channel, voltage gated eag related subfamily H, member 6
KCNK15 potassium channel, two pore domain subfamily K, member 15
KDM5A lysine (K)-specific demethylase 5A
KHK ketohexokinase (fructokinase)
KIAA0232 KIAA0232
KIAA1456 KIAA1456
KIDINS220 kinase D-interacting substrate, 220kDa
KIT v-kit Hardy-Zuckerman 4 feline sarcoma viral oncogene homolog
KLF11 Kruppel-like factor 11
KLHL29 kelch-like family member 29
KLHL42 kelch-like family member 42
KLHL5 kelch-like family member 5
KLHL8 kelch-like family member 8
KLRAP1 killer cell lectin-like receptor subfamily A pseudogene 1
KRAS Kirsten rat sarcoma viral oncogene homolog
KRBA2 KRAB-A domain containing 2
KREMEN1 kringle containing transmembrane protein 1
KRT36 keratin 36, type I
KRT76 keratin 76, type II
KRT84 keratin 84, type II
KRTAP21-1 keratin associated protein 21-1
LAMTOR3 late endosomal/lysosomal adaptor, MAPK and MTOR activator 3
LBH limb bud and heart development
LCLAT1 lysocardiolipin acyltransferase 1
LCORL ligand dependent nuclear receptor corepressor-like
LCP2 lymphocyte cytosolic protein 2 (SH2 domain containing leukocyte protein of 76kDa)
LEMD3 LEM domain containing 3
LILRA1 leukocyte immunoglobulin-like receptor, subfamily A (with TM domain), member 1
LILRB1 leukocyte immunoglobulin-like receptor, subfamily B (with TM and ITIM domains), member 1
LILRB2 leukocyte immunoglobulin-like receptor, subfamily B (with TM and ITIM domains), member 2
LILRP2 leukocyte immunoglobulin-like receptor pseudogene 2
LIN54 lin-54 DREAM MuvB core complex component
LINC00028 long intergenic non-protein coding RNA 28
LINC00159 long intergenic non-protein coding RNA 159
LOC340074 uncharacterized LOC340074
LOC642846 DEAD/H (Asp-Glu-Ala-Asp/His) box polypeptide 11-like
LOC644145 exocyst complex component 1 pseudogene
LOH12CR1 loss of heterozygosity, 12, chromosomal region 1
LPCAT3 lysophosphatidylcholine acyltransferase 3
LRGUK leucine-rich repeats and guanylate kinase domain containing
LRP1 low density lipoprotein receptor-related protein 1
LRP6 low density lipoprotein receptor-related protein 6
LRRC23 leucine rich repeat containing 23
LRRC37A3 leucine rich repeat containing 37, member A3
LRRC37BP1 leucine rich repeat containing 37B pseudogene 1
LRRC66 leucine rich repeat containing 66
LRRC75A leucine rich repeat containing 75A
LRRTM1 leucine rich repeat transmembrane neuronal 1
LTBR lymphotoxin beta receptor (TNFR superfamily, member 3)
MAGOHB mago-nashi homolog B (Drosophila)
MAGT1 magnesium transporter 1
MANBA mannosidase, beta A, lysosomal
MAP9 microtubule-associated protein 9
MAPRE3 microtubule-associated protein, RP/EB family, member 3
MED21 mediator complex subunit 21
MEDAG mesenteric estrogen-dependent adipogenesis
MEIS3P1 Meis homeobox 3 pseudogene 1
MFAP3L microfibrillar-associated protein 3-like
MGAT3 mannosyl (beta-1,4-)-glycoprotein beta-1,4-N-acetylglucosaminyltransferase
MIER3 mesoderm induction early response 1, family member 3
MLF2 myeloid leukemia factor 2
MOB1B MOB kinase activator 1B
MRPL51 mitochondrial ribosomal protein L51
MRPS35 mitochondrial ribosomal protein S35
MSRB3 methionine sulfoxide reductase B3
MTCH1 mitochondrial carrier 1
MYL10 myosin, light chain 10, regulatory
NAMPT nicotinamide phosphoribosyltransferase
NBPF3 neuroblastoma breakpoint family, member 3
NBPF7 neuroblastoma breakpoint family, member 7
NCAM1 neural cell adhesion molecule 1
NCAPD2 non-SMC condensin I complex, subunit D2
NCOA1 nuclear receptor coactivator 1
NDUFA9 NADH dehydrogenase (ubiquinone) 1 alpha subcomplex, 9, 39kDa
NECAP1 NECAP endocytosis associated 1
NEGR1 neuronal growth regulator 1
NFASC neurofascin
NFATC2 nuclear factor of activated T-cells, cytoplasmic, calcineurin-dependent 2
NFXL1 nuclear transcription factor, X-box binding-like 1
NHEG1 neuroblastoma highly expressed 1
NHLRC3 NHL repeat containing 3
NINJ2 ninjurin 2
NOL10 nucleolar protein 10
NTN3 netrin 3
NXF4 nuclear RNA export factor 4 pseudogene
OR13D1 olfactory receptor, family 13, subfamily D, member 1
OR2D2 olfactory receptor, family 2, subfamily D, member 2
OR4C3 olfactory receptor, family 4, subfamily C, member 3
OR51E1 olfactory receptor, family 51, subfamily E, member 1
OR52E2 olfactory receptor, family 52, subfamily E, member 2
OR52W1 olfactory receptor, family 52, subfamily W, member 1
P3H2 prolyl 3-hydroxylase 2
P3H3 prolyl 3-hydroxylase 3
PAFAH1B1 platelet-activating factor acetylhydrolase 1b, regulatory subunit 1 (45kDa)
PCDH7 protocadherin 7
PCDHA5 protocadherin alpha 5
PCDHA8 protocadherin alpha 8
PCDHGB6 protocadherin gamma subfamily B, 6
PCGF3 polycomb group ring finger 3
PCSK6 proprotein convertase subtilisin/kexin type 6
PDIA6 protein disulfide isomerase family A, member 6
PEX1 peroxisomal biogenesis factor 1
PEX5 peroxisomal biogenesis factor 5
PGAP1 post-GPI attachment to proteins 1
PHIP pleckstrin homology domain interacting protein
PHKA1 phosphorylase kinase, alpha 1 (muscle)
PHOX2A paired-like homeobox 2a
PKD2 polycystic kidney disease 2 (autosomal dominant)
PLA2G2A phospholipase A2, group IIA (platelets, synovial fluid)
PLAG1 pleiomorphic adenoma gene 1
PLAT plasminogen activator, tissue
PLBD1 phospholipase B domain containing 1
POLR2B polymerase (RNA) II (DNA directed) polypeptide B, 140kDa
POU5F1B POU class 5 homeobox 1B
PPFIBP1 PTPRF interacting protein, binding protein 1 (liprin beta 1)
PPIL1 peptidylprolyl isomerase (cyclophilin)-like 1
PPP1CB protein phosphatase 1, catalytic subunit, beta isozyme
PPP1R12B protein phosphatase 1, regulatory subunit 12B
PPP1R9A protein phosphatase 1, regulatory subunit 9A
PQLC3 PQ loop repeat containing 3
PREB prolactin regulatory element binding
PRH1 proline-rich protein HaeIII subfamily 1
PRKAA2 protein kinase, AMP-activated, alpha 2 catalytic subunit
PRKCA protein kinase C, alpha
PRSS35 protease, serine, 35
PRSS42 protease, serine, 42
PTMS parathymosin
PTPRZ1 protein tyrosine phosphatase, receptor-type, Z polypeptide 1
PUM2 pumilio RNA-binding family member 2
PYROXD1 pyridine nucleotide-disulphide oxidoreductase domain 1
RAB10 RAB10, member RAS oncogene family
RAD52 RAD52 homolog (S. cerevisiae)
RAMP3 receptor (G protein-coupled) activity modifying protein 3
RBMS1 RNA binding motif, single stranded interacting protein 1
RBPJ recombination signal binding protein for immunoglobulin kappa J region
RDH14 retinol dehydrogenase 14 (all-trans/9-cis/11-cis)
RECQL RecQ helicase-like
REP15 RAB15 effector protein
RFX2 regulatory factor X, 2 (influences HLA class II expression)
RHNO1 RAD9-HUS1-RAD1 interacting nuclear orphan 1
RNASE11 ribonuclease, RNase A family, 11 (non-active)
RND3 Rho family GTPase 3
RNF113B ring finger protein 113B
RNF8 ring finger protein 8, E3 ubiquitin protein ligase
ROCK2 Rho-associated, coiled-coil containing protein kinase 2
RPL13P5 ribosomal protein L13 pseudogene 5
RUNX1 runt-related transcription factor 1
SCARB2 scavenger receptor class B, member 2
SDAD1 SDA1 domain containing 1
SEMA3E sema domain, immunoglobulin domain (Ig), short basic domain, secreted, (semaphorin) 3E
SEPT11 septin 11
SERPINB11 serpin peptidase inhibitor, clade B (ovalbumin), member 11 (gene/pseudogene)
SH3BP2 SH3-domain binding protein 2
SKIV2L2 superkiller viralicidic activity 2-like 2 (S. cerevisiae)
SLC16A4 solute carrier family 16, member 4
SLC2A14 solute carrier family 2 (facilitated glucose transporter), member 14
SLC2A3 solute carrier family 2 (facilitated glucose transporter), member 3
SLC30A10 solute carrier family 30, member 10
SLC30A6 solute carrier family 30 (zinc transporter), member 6
SLC35B4 solute carrier family 35 (UDP-xylose/UDP-N-acetylglucosamine transporter), member B4
SLC38A9 solute carrier family 38, member 9
SLC4A1AP solute carrier family 4 (anion exchanger), member 1, adaptor protein
SLC52A1 solute carrier family 52 (riboflavin transporter), member 1
SLC5A6 solute carrier family 5 (sodium/multivitamin and iodide cotransporter), member 6
SMIM10L2A small integral membrane protein 10 like 2A
SMIM20 small integral membrane protein 20
SNORA45B small nucleolar RNA, H/ACA box 45B
SNORA71C small nucleolar RNA, H/ACA box 71C
SNX10 sorting nexin 10
SNX18 sorting nexin 18
SNX31 sorting nexin 31
SPAST spastin
SPATA22 spermatogenesis associated 22
SPATS1 spermatogenesis associated, serine-rich 1
SPCS3 signal peptidase complex subunit 3 homolog (S. cerevisiae)
SPTBN1 spectrin, beta, non-erythrocytic 1
SSUH2 ssu-2 homolog (C. elegans)
ST8SIA1 ST8 alpha-N-acetyl-neuraminide alpha-2,8-sialyltransferase 1
STBD1 starch binding domain 1
STK17A serine/threonine kinase 17a
STK32B serine/threonine kinase 32B
STK38L serine/threonine kinase 38 like
STRAP serine/threonine kinase receptor associated protein
STRIP2 striatin interacting protein 2
SUPT7L suppressor of Ty 7 (S. cerevisiae)-like
TAAR6 trace amine associated receptor 6
TAPBPL TAP binding protein-like
TAS2R13 taste receptor, type 2, member 13
TAS2R14 taste receptor, type 2, member 14
TAS2R19 taste receptor, type 2, member 19
TAS2R20 taste receptor, type 2, member 20
TAS2R31 taste receptor, type 2, member 31
TAS2R50 taste receptor, type 2, member 50
TBC1D22B TBC1 domain family, member 22B
TBC1D28 TBC1 domain family, member 28
TBC1D29 TBC1 domain family, member 29
TBK1 TANK-binding kinase 1
TBX15 T-box 15
TC2N tandem C2 domains, nuclear
TEC tec protein tyrosine kinase
THAP5 THAP domain containing 5
THAP6 THAP domain containing 6
TIGAR TP53 induced glycolysis regulatory phosphatase
TM7SF3 transmembrane 7 superfamily member 3
TMEM106A transmembrane protein 106A
TMEM128 transmembrane protein 128
TMEM150C transmembrane protein 150C
TMEM192 transmembrane protein 192
TMEM217 transmembrane protein 217
TMPRSS11B transmembrane protease, serine 11B
TMTC1 transmembrane and tetratricopeptide repeat containing 1
TMX4 thioredoxin-related transmembrane protein 4
TNFRSF1A tumor necrosis factor receptor superfamily, member 1A
TNP1 transition protein 1 (during histone to protamine replacement)
TNS3 tensin 3
TRAF1 TNF receptor-associated factor 1
TRIM56 tripartite motif containing 56
TRPC1 transient receptor potential cation channel, subfamily C, member 1
TRPM8 transient receptor potential cation channel, subfamily M, member 8
TSHZ2 teashirt zinc finger homeobox 2
TSPAN11 tetraspanin 11
TTC27 tetratricopeptide repeat domain 27
TTLL6 tubulin tyrosine ligase-like family member 6
TYW1B tRNA-yW synthesizing protein 1 homolog B (S. cerevisiae)
UBA6 ubiquitin-like modifier activating enzyme 6
UBXN2A UBX domain protein 2A
UGDH UDP-glucose 6-dehydrogenase
UGT2B7 UDP glucuronosyltransferase 2 family, polypeptide B7
UPP2 uridine phosphorylase 2
USO1 USO1 vesicle transport factor
USP5 ubiquitin specific peptidase 5 (isopeptidase T)
UVSSA UV-stimulated scaffold protein A
VSTM2L V-set and transmembrane domain containing 2 like
VWA5A von Willebrand factor A domain containing 5A
WBP11 WW domain binding protein 11
WDFY3 WD repeat and FYVE domain containing 3
WDR19 WD repeat domain 19
WIPF1 WAS/WASL interacting protein family, member 1
WNK1 WNK lysine deficient protein kinase 1
XPOT exportin, tRNA
YARS2 tyrosyl-tRNA synthetase 2, mitochondrial
YIPF4 Yip1 domain family, member 4
YPEL5 yippee-like 5 (Drosophila)
YWHAQ tyrosine 3-monooxygenase/tryptophan 5-monooxygenase activation protein, theta
ZACN zinc activated ligand-gated ion channel
ZBTB49 zinc finger and BTB domain containing 49
ZC3H6 zinc finger CCCH-type containing 6
ZCCHC4 zinc finger, CCHC domain containing 4
ZDHHC2 zinc finger, DHHC-type containing 2
ZFAND3 zinc finger, AN1-type domain 3
ZNF141 zinc finger protein 141
ZNF287 zinc finger protein 287
ZNF384 zinc finger protein 384
ZNF491 zinc finger protein 491
ZNF512 zinc finger protein 512
ZNF547 zinc finger protein 547
ZNF721 zinc finger protein 721
ZNF805 zinc finger protein 805
ZNF862 zinc finger protein 862
ZNF878 zinc finger protein 878
ZSCAN12P1 zinc finger and SCAN domain containing 12 pseudogene 1

low expression

Symbol Name
EIF2B5 eukaryotic translation initiation factor 2B, subunit 5 epsilon, 82kDa